Lily is the leader of the Resistance in NieR:Automata Ver1.1a. She has a steadfast personality and cares for her friends. She first appears in the novella YoRHa ̶Ver. 1.05, which tells the events of the YoRHa stage play. The manga YoRHa: Pearl Harbor Descent Record also tells the events from the stage play. Originally, she became part of the Resistance under the command of Rose during the 8th Machine War that took place over 200 years before Ver1.1a.