Papillon Rose is the title character, leader of the Papillon Soldiers and the secret identity of Tsubomi. She was the first Papillon Soldier to awaken. Tsubomi is working in the lingerie pub called 'Papillon'. While running to work from school, Tsubomi bumps into Hikaru Shishio, and proceeds to having sex with him in a love motel. The next night at the Papillon Pub, she meets a black talking cat named Rama. She tells her her name is Rama and that she is searching for the Papillon Soldiers. At that moment, the pub "Koneko clubhouse" is attacked by a woman named Sister Biene. Tsubomi is trapped by Biene and her Secret shrine starts to glow, revealing her as Papillon Rose. Months after Papillon Rose and her fellow soldiers defeated Regina Apis, they spend their nights doing nothing more then sex. Tsubomi stays in an FWB relationship with Razy Minowa, who she sleeps with wearing her battle lingerie. Meanwhile, a trans organisation named Gel Dynasty run by Shiorinite is planning to spread an STD virus using a weapon. Papillon Rose, Margarette and Lily are surrounded by a group of transvestites, and Rose gets influenced by the virus. Recovering from the illness, Rose's powers are improved, so she defeats the enemy group. Later, Papillon Rose is attacked by her comrades after being influenced by the food virus. She recieves the ultimate humiliation made to recover the soldiers from the virus's influence and joins the other soldiers against the black snake leader, Hydra, who plans to release a lust virus on planet earth.
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